can not work. It is probable that many monoliths and some old walls have fallen down from having been undermined by worms. THEY PREPARE THE GROUND FOR SEED. Worms prepare the ground in an excellent Page 309. manner for the growth of fibrous-rooted plants and for seedlings of all kinds. They periodically expose the mold to the air, and sift it so that no stones larger than the particles which they can swallow are left in it. They mingle the whole intimately together, like a gardener who prepares fine soil for his choicest plants. In this state it is well fitted to retain moisture and to absorb all soluble substances, as well as for the process of nitrification. The bones of dead animals, the harder parts of insects, the shells of land-mollusks, leaves, twigs, etc., are before long all buried beneath the accumulating castings of worms, and are thus brought in a more or less decayed state within reach of the roots of plants. Worms likewise drag an infinite number of dead leaves and other parts of plants into their burrows, partly for the sake of plugging them up and partly as food. The leaves which are dragged into the burrows as food, after being torn into the finest shreds, partially digested, and saturated with the intestinal and urinary secretions, are commingled with much earth. This earth forms the dark-colored, rich humus which almost everywhere covers the surface of the land with a fairly welldefined layer or mantle. Von Hensen placed two worms in a vessel eighteen inches in diameter, which was filled with sand, on which fallen leaves were strewed; and these were soon dragged into their burrows to a depth of three inches. After about six weeks an almost uniform layer of sand, a centimetre (4 inch) in thickness, was converted into humus by having passed through the alimentary canals of these two worms. It is believed by some persons that worm-burrows, which often penetrate the ground almost perpendicularly to a depth of five or six feet, materially aid in its drainage; notwithstanding that the viscid castings piled over the mouths of the burrows prevent or check the rain-water directly entering them. They allow the air to penetrate deeply into the ground. They also greatly facilitate the downward passage of roots of moderate size; and these will be nourished by the humus with which the burrows are lined. Many seeds owe their germination to having been covered by castings; and others buried to a considerable depth beneath accumulated castings lie dormant, until at some future time they are accidentally uncovered and germinate. When we behold a wide, turf-covered exPage 313. panse, we should remember that its smoothness, on which so much of its beauty depends, is mainly due to all the inequalities having been slowly leveled by worms. It is a marvelous reflection that the whole of the superficial mold over any such expanse has passed, and will again pass, every few years through the bodies of worms. The plow is one of the most ancient and most valuable of man's inventions; but long before he existed the land was in fact regularly plowed, and still continues to be thus plowed, by earth-worms. It may be doubted whether there are many other animals which have played so important a part in the history of the world as have these lowly organized creatures. Some other animals, however, still more lowly organized, namely corals, have done far more conspicuous work in having constructed innumerable reefs and islands in the great oceans; but these are almost confined to the tropical zones. INTELLIGENCE OF WORMS. We can hardly escape from the conclusion Page 91. that worms show some degree of intelligence in their manner of plugging up their burrows. Each particular object is seized in too uniform a manner, and from causes which we can generally understand, for the result to be attributed to mere chance. That every object has not been drawn in by its pointed end, may be accounted for by labor having been saved through some being inserted by their broader or thicker ends. No doubt worms are led by instinct to plug up their burrows; and it might have been expected that they would have been led by instinct how best to act in each particular case, independently of intelligence. We see how difficult it is to judge whether intelligence comes into play, for even plants might sometimes be thought to be thus directed; for instance, when displaced leaves redirect their upper surfaces toward the light by extremely complicated movements and by the shortest course. With animals, actions appearing due to intelligence may be performed through inherited habit without any intelligence, although aboriginally thus acquired. Or the habit may have been acquired through the preservation and inheritance of beneficial variations of some other habit; and in this case the new habit will have been acquired independently of intelligence throughout the whole course of its development. There is no a priori improbability in worms having acquired special instincts through either of these two latter means. Nevertheless, it is incredible that instincts should have been developed in reference to objects, such as the leaves or petioles of foreign plants, wholly unknown to the progenitors of the worms which act in the described manner. Nor are their actions so unvarying or inevitable as are most true instincts. As worms are not guided by special instincts in each particular case, though possessing a general instinct to plug up their burrows, and, as chance is excluded, the next most probable conclusion seems to be that they try in many different ways to draw in objects, and at last succeed in some one way. But it is surprising that an animal so low in the scale as a worm should have the capacity for acting in this manner, as many higher animals have no such capacity. Mr. Romanes, who has specially studied the Page 95. minds of animals, believes that we can safely infer intelligence only when we see an individual profiting by its own experience. Now, if worms try to drag objects into their burrows first in one way and then in another, until they at last succeed, they profit at least in each particular instance by experience. One alternative alone is left, namely, that Page 98. worms, although standing low in the scale of organization, possess some degree of intelligence. This will strike every one as very improbable; but it may be doubted whether we know enough about the nervous system of the lower animals to justify our natural distrust of such a conclusion. With respect to the small size of the cerebral ganglia, we should remember what a mass of inherited knowledge, with some power of adapting means to an end, is crowded into the minute brain of a workerant. III. THE LAWS OF VARIABILITY WITH RESPECT The Variation of Animals and Domestica page 3. ΤΟ ΑΝΙMALS AND PLANTS. I SHALL in this volume treat, as fully as my materials permit, the whole subject of variaPlants under tion under domestication. We may thus hope tion, vol. i, to obtain some light, little though it be, on the causes of variability, on the laws which govern it-such as the direct action of climate and food, the effects of use and disuse, and of correlation of growthand on the amount of change to which domesticated organisms are liable. Although man does not cause variability and can not even prevent it, he can select, preserve, and accumulate the variations given to him by the hand of Nature almost in any way which he chooses; and thus he can certainly produce a great result. Selection may be followed either methodically and intentionally, or unconsciously and unintentionally. Man may select and preserve each successive variation, with the distinct intention of improving and altering a breed, in accordance with a preconceived idea; and by thus adding up variations, often so slight as to be imperceptible by an uneducated eye, he has effected wonderful changes and improvements. It can, also, be clearly shown that man, without any intention or thought |